The Biology Ph.D. Program is divided into four subprograms that center on separate, but overlapping, areas of biology. Learn about the differences in curriculum associated with each subprogram (and about any specific application requirements associated with each).

Ecology, Evolutionary Biology, and Behavior (EEB)

a yellow african weaverbird perched on the underside of its woven spherical nest, holding a frilly branch in its beak

Behavior, ecology, evolution, and systematics, from anatomy to zoogeography. Faculty include members of the curatorial staff of the American Museum of Natural History.

Learn more about the EEB Subprogram »

Molecular, Cellular, and Developmental Biology (MCD)

microscope image of cell structure

Molecular basis of cell function, of prokaryotic systems, of animal and plant development, and of complex biological processes such as the immune and nervous systems.

Learn more about the MCD Subprogram »

Neuroscience (NS)

digital rendering of neurons in blue on a black background

Organization and function of nervous systems, approached from biochemical, cellular, organismal, and environmental perspectives. Faculty include staff of the New York State Institute for Basic Research in Developmental Disabilities.

Learn more about the NS subprogram »

Plant Sciences (PS)

tight focus photo of small green seedlings germinating in a petri dish held between the thumb and forefinger of a gloved hand

Study of major plant taxa from bryophytes to flowering plants at every level of organization from the molecular to the ecosystem. Work ranges from theoretical to socioeconomic. Faculty include members of the curatorial staff of the New York Botanical Garden.

Learn more about the PS subprogram »

a cloudy sky reflected in the surface of a calm lake, a small strip of rocky beach in the foreground, ringed by trees in early autumn colors

Learn, explore, and share ideas in the Poconos

Subprogram Retreats

Each year, Biology program faculty, students and guests have the opportunity to gather with the members of their subprograms for a rejuvenating retreat at the Pocono Environmental Education Center (PEEC) in Dingman’s Ferry, Pennsylvania.

Learn more about the annual retreats