- Neurophysiology and neuroanatomy of the central visual pathways, organization and development of mammalian cerebral cortex
- Senior Research Fellow, Institute of Ophthalmology, University College London, 1992-1998
- Postdoctoral Fellow, University of Pittsburgh School of Medicine, 1990-1992 M.A.
- Ph.D. in Experimental Psychology, Center for Neural Science, NYU, 1990
- B.A. in Biophysics, University of Pennsylvania,1984